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Accurate seismic images of the crust are essential for assessing seismic hazards and elucidating tectonic processes that shape surface landforms. Although California and Nevada have been studied extensively using various seismic datasets and tomographic methods, the region lacks a seismic model that can accurately define both the shallow (<8 km) and deeper crust. We take the advantage of recent increases in seismic data coverage to build a new 3D shear wave speed model by jointly inverting Rayleigh wave ellipticity, phase velocity, and teleseismic P waveforms. In the Great Valley, the new model reveals an asymmetric basement, steeply dipping in the west and gently dipping in the east. Beneath its western margin, in the Coast Ranges, we resolve a wedge‐shaped, low‐velocity zone in the upper‐middle crust, interpreted as Franciscan Complex. Our images confirm that uplift of the western Great Valley and an eastward shift of its depositional center are caused by wedging and underthrusting of the complex during subduction. Across the Basin and Range, the resolved crust has an average thickness of 38 km in the southern half of the northern Basin and Range, about 5 km thicker than neighboring regions. The thickened crust overlaps with major volcanic centers of the mid‐Cenozoic ignimbrite flare‐up. This spatial correlation may suggest magmatic intrusions and underplating contributed to crustal growth and thickening prior to Miocene Basin and Range extension. Overall, the new model is consistent with active source studies in the region but provides a more comprehensive view of shallow and deep structures across this large and tectonically complex region.